PhD fellowships at University of Copenhagen, Denmark

16 Best PhD fellowships at University of Copenhagen, Denmark

Online applications are being accepted by the University of Copenhagen in Denmark for a variety of fully funded PhD fellowships in various Departments. Here is a list of PhD programmes at the University of Copenhagen in Denmark that are fully funded. Any qualified candidate is encouraged to apply right away.

(1) Marie Sklodowska-Curie PhD fellowship: Cellular memory and intestinal regeneration

We are seeking an ambitious, highly motivated, and technically skilled candidate preferably with experience in cell and molecular biology. The goal of the project is to decipher the cellular and molecular responses following tissue damage and during tissue regeneration in the intestinal epithelium using mouse models and patient samples. The intestinal epithelium has an amazing capacity to regenerate itself following damage via processes that involve pronounced changes in the cellular identity (see e.g. Larsen and Jensen, 2019, Curr Opin Genet Dev). We have demonstrated that this process depends on changes in the environment leading to integrin-based activation of mechanosensory pathway driven dedifferentiation processes vital for tissue regeneration (see Yui et al., 2018, Cell Stem Cell; Schweiger et al., 2022, Gastroenterology). Yet, how these changes impact cellular memory and what are the long-term consequences on cell fate decisions remain largely unknown. In this project, we will take advantage of established and optimised tools including single cell omics, lineage tracing and organoid technologies to address whether and how injury and the associated dedifferentiation influence cell behavior long-term.

The REGENERATE-IT Consortium

This project is a part of the REGENERATE-IT Doctoral Network, funded by the Marie-Sklodowska-Curie Action (MSCA). The network brings together a full spectrum of disciplines – from fundamental to applied research – to train the next generation of researchers in regenerative biology and adjacent fields. It is a consortium of high-profile researchers from various universities, research institutions, and companies spanning 12 academic institutions in 10 European countries (the Netherlands, France, Belgium, Austria, Sweden, Germany, Italy, Denmark, Switzerland, and the UK). The competencies available in the REGENERATE-IT Network are further expanded by the involvement of one non-academic beneficiary, four industrial partner organisations, a patent attorney firm, a non-profit research organisation, a scientific publisher, a patient advocacy group, and Women Scientists network organisation.

The REGENERATE-IT consortium offers 12 fully funded PhD positions to work with leading European researchers to advance the frontiers of regenerative biology and medicine. More information about all projects is available on the project website

Deadline : 31 March 2023

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 (2) 8 fully funded Marie S. Curie PhD positions at the Biotech Research and Innovation Centre (BRIC)

Up to 8 PhD positions are open in the international Marie S. Curie Doctoral Programme ‘Discovering treatment from biomedical research’ (DISCOVER). The positions are open for talented young researchers with a master’s degree, who want to pursue a PhD degree in biomedical research. The programme is co-funded by the European Union’s Horizon 2020 Research and Innovation Programme under the Marie Skłodowska-Curie grant agreement no. 101034291.

About the programme

DISCOVER will in total recruit 24 fellows over three calls during 2021-2023. The programme offers 36-months fully funded positions, attractive employment and salary conditions with enrolment as PhD student at the Graduate School of Health and Medical Sciences, UCPH. Each fellow will work with an original biomedical research project in one of BRIC’s laboratories. Clinical co-supervision will ensure to develop the translational potential of the project. An individual career development plan containing research and transferable courses, international research visits, mentoring and career activities, is also a central element. Please visit for more information on the programme.

Deadline : 25 March 2023

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(3) PhD position: Physics of Life

The project is focused on the viruses infecting bacteria (bacteiophages): they are the drivers of microbial ecology, gene transfer and microbial evolution, and also an ideal in-vivo model system to study the interplay between a virus and its host. We aim to explore the virus development upon infection as a dynamic and stochastic process exhibiting decisions. We are also actively studying the role of phages at the population scale and ecology scale and as pattern formation, to the macroscopic scale of epidemics as it spread in a 2 or 3-dimensional geometry. We have a strong connection between experimental and theoretical research: We develop mathematical and computational models to mimic, suggest and perform experiments on these fascinating systems.

Who are we looking for?

We are looking for a talented, highly motivated and creative scientist who is interested in exploring fundamental questions in the exciting field of the physics of life. Scientists with strong mathematical and computational skills in physics/complex systems who want to extend their skills to biological systems with close collaboration with experimental biology are strongly encouraged to apply. The successful candidates must be imaginative and have the curiosity to drive the project. The candidate must have excellent English skills, both written and oral, and hold a Master’s degree in physics, biophysics, quantitative biology, complex systems, or related disciplines. If you are (or are eligible to be) enrolled at one of the faculty’s master programs in physics, it is also an option to be enrolled in the integrated MSc and PhD program

Deadline : 30 April 2023

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(4) PhD fellowship: Fate of Nanofertilisers in Soils

The PhD project is part of a larger Novo Nordisk Fonden research project called BioComFertBiocompatible nanofertilizers for targeted delivery and programmed release of essential mineral ions in crops, led by Professor Søren Husted at PLEN, UCPH and involving several research groups at UCPH and DTU (DK) and at McGill University (Canada). The overall aim of BioComFert is to develop novel nanomaterials for targeted delivery of nutrients via micro- and nanoporous leaf cuticles to the specific plant tissue and organelles where they are needed. A set of nanomaterials are currently being developed with different nutrient loads (i.e., P, Mn) and surface properties, and their effectiveness tested on different plants, involving nanoscale characterisation tools to monitor entry and delivery pathways of these nanofertilisers, and plant nutritional status.

The aim of this PhD project will be to assess the environmental fate of these nanofertilisers when they end up in the soil. It is very likely that when nanofertilisers are sprayed onto crops, a fraction will enter the soil directly, and others may enter later, e.g., during rainfall/wash-off, or during other climate stresses. Thus, before these newly developed nanofertilisers (NPs) can be applied to crops for real, it is critical to monitor their fate in soil and assess potential losses into the environment. This will include NP mobility studies in model and field test soils, under varied environmental and climate stresses, as well as assessment of NP stability, i.e., dissolution and transformation behaviour, and their ultimate impact on soil biogeochemistry. In BioComFert, a variety of state of the art characterisation tools are being employed and methods develop (e.g., X-ray micro-CT, fluorescence and electron microscopy), and the candidate will be encouraged to make use of these tools and further develop protocols to monitor the fate of NPs in the soils.

Who are we looking for?
We are looking for a highly dedicated and enthusiastic candidate with interest in soil biogeochemistry and nanomaterials and background in environmental sciences, soil sciences, geochemistry or a related field. The project involves extensive laboratory work and characterisation of solids and solutions using various standard but also high-end characterisation tools, so a keen interest for learning and applying state of the art equipment and developing robust laboratory workflow procedures is a must. This project is closely interlinked with other BioComFert PhD and Postdoc fellowships, thus the candidate is expected to interact and communicate regularly with the other fellows and PIs in the project, to report on new results and participate in project discussions.

Deadline :30 April 2023

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(5) PhD scholarship: Life-time imaging of the chemical microenvironment and metabolic activity in biofilms and 3D bioprinted constructs

The scientific objective of FLIMagin3D is to synergistically evolve key aspects of Fluorescence Lifetime Imaging Microscopy (FLIM) across disciplines by i) advancing the acquisition of FLIM data and the analysis pipeline, ii) developing improved platforms that facilitate imaging for a wide range of (bio)medical, chemical and biophysical applications, and iii) advancing the state-of-the-art in FLIM biosensors. The project is a collaboration between 8 academic and 4 industrial partners from 7 European countries and brings together highly synergistic expertise on advanced bioimaging and image analysis.

The specific PhD project will be within the Microenvironmental Ecology &Symbiosis research group headed by Prof Michael Kühl at the Department of Biology, University of Copenhagen with the main workplace at the Marine Biological Section in Helsingør ( Besides in-house experiments on time- and frequency-domain lifetime imaging platforms, the studies will also involve work with microscopy platforms at the Center of Advanced Bioimaging ( in Copenhagen. The group has pioneered microenvironmental analyses and chemical imaging of natural aquatic systems and organisms, as well as chemical microenvironments in 3D bioprinted constructs. 

Deadline : 15 April 2023

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(6) PhD fellowship: Image analysis using deep learning at Department of Drug Design and Pharmacology

We are seeking a highly motivated PhD student to work on this joint project between the Eye Translational Research Unit at The University of Copenhagen and the Section for Visual Computing at DTU Compute. The project, called “Finding Ophthalmic Risk and Evaluating the Value of Eye exams and their Predictive Reliability – Optical Coherence Tomography” (FOREVER-OCT), is a subset of the ongoing FOREVER project ( The PhD project is focused on identifying risk factors for eye diseases based on fundus photographs and optical coherence tomography (OCT) scans of the participants in the FOREVER project.

In the FOREVER project, 30,000 fundus photographs and 10,000 OCT scans are collected for the FOREVER cohort. Fundus photographs are widely used in Denmark, but OCT is a relatively new imaging examination and not as widely incorporated in an optometric setting. During this PhD project, the research will be focused on:

  • Investigate the diagnostic value of the OCT imaging modality in the diagnosis of ophthalmic diseases, compared to fundus images.
  • Investigate the possibilities of a combined diagnostic model based on both fundus and OCT scans, and possibly other clinical data.
  • Evaluate potential associations between OCT parameters and risk factors for glaucoma.

As a PhD student on this project, you will have the opportunity to work with a team of experienced researchers, clinicians, and data scientists on tasks including data collection and analysis. Your role will involve discussing and applying state-of-the-art deep learning methods to the FOREVER dataset. You will also be responsible for carrying out the analyses and assessing the results in a clinical context. As this is a joint project, it is expected that you will divide your time between the two different research environments at DTU Compute and the University of Copenhagen. You will publish your findings in academic journals and present your work at international conferences.

Deadline : 1 April 2023

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(7) PhD fellowships:  Chemical Biology at the Department of Drug Design and Pharmacology

PhD project 1Development of peptide-based probes to modulated biomolecular condensates at PSD
In this project, the fellow will examine all PPIs of the PSD and delineate their peptide binding epitopes. The identified binding epitopes will be further optimized and (peptide therapeutic maturation) evaluated in in vitro biochemical and LLPS assays. Multiple rounds of affinity maturation will yield peptides either disrupting or stabilizing the relevant PPIs and restoring a physiological PSD. Ultimately, this will provide a better understanding how to restore the healthy state of PSD using peptide modulators.

PhD project 2Elucidating aberrant biomolecular condensates in the PSD
The five key proteins present in the PSD (PSD-95, SynGAP, Homer, Shank and GKAP) are important for normal brain function, and dysfunction of these proteins has vast implications for a wide range of brain diseases. In this project the fellow will first explore the correlation between brain disorders and biomolecular condensates by introducing specific disease-relevant patient mutations into the proteins of interest and observe the effects on LLPS.  Additionally, the resulting impact of disease mutations on LLPS-PSD will guide efforts towards developing compounds to tackle neurological disorders.

PhD project 3Development of CNS-permeant peptide antivirals
The first part of the project involves developing peptides inhibiting infectivity by disrupting the spike protein receptor binding domain interactions with the human angiotensin-converting enzyme 2 receptor, the binding target of the viral spike protein. The second involves developing peptides inhibiting virulence by targeting interactions of the viral envelope protein with host proteins containing PDZ domains.

Deadline : 1 April 2023

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(8) PhD position: Medical Doctor (læge) for a PhD Fellowship in Experimental Gut Physiology at the Department of Biomedical Sciences

The position is available in the Experimental Gut Physiology Group at the Department of Biomedical Sciences (BMI) headed by assistant professor/specialist registrar Simon Veedfald, who will be your supervisor in collaboration with professors Jens Juul Holst (Department of Biomedical Sciences) and Sten Madsbad (Department of Endocrinology, Hvidovre Hospital). The stated goal of the Department of Biomedical Sciences is to pursue basic biomedical research with an emphasis on clinical translation. We are working towards a better understanding of the physiology of the gut and towards identifying new drug candidates to improve the treatment of intestinal and extra intestinal disease.

Project description

As the chosen PhD candidate you will be responsible for recruiting and conducting clinical experimental work on a daily basis, collecting data, performing data analyses and disseminating the study results in manuscripts and giving presentations. You will principally perform clinical experimental work at the clinical research facility at Hvidovre Hospital. You will be working alongside clinical researchers working on a number of different studies in a collaborative and supportive environment.

At BMI you will learn and perform peptide hormone analyses and be introduced to the other experimental approaches used in the laboratory of professor Jens Juul Holst.

As part of your PhD fellowship, you will take relevant PhD courses and participate in local and international scientific meetings. You will become familiar with teaching methodology and be involved in the teaching of medical students.

Deadline : 31 March 2023

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(9) PhD fellowship in Particle Astrophysics

This project aims at solving one of the most urgent riddles in particle astrophysics: how neutrinos affect the physics of the death of massive stars as core-collapse supernova explosions and the merger of two neutron stars or a neutron star and a black hole. Neutrinos are feebly interacting particles copiously produced in these dense sources. Neutrinos exist in three different kinds, or flavors, and have the fascinating property of changing their flavor while propagating (flavor conversion). Because of the high density of neutrinos in the core of supernovae or compact binary mergers, flavor conversion becomes a non-linear phenomenon, whose understanding is quite preliminary. Numerical simulations buttressed by analytic diagnostic methods will be employed to radically advance our understanding within a multi-messenger framework. This project will have implications on fundamental physics, the origin of the heavy elements, as well as our comprehension of the behavior of matter at extreme densities and the physics of  neutrino-dense sources.

Deadline : 16 April 2023

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(10) PhD stipends in Statistics, and the mathematics of Insurance and Economics

Department of Mathematical Sciences (MATH) invites applications for PhD stipends in applied mathematics, including statistics, the mathematics of insurance and economics, and mathematical biology. Start date is (expected to be) September 2023 or as soon as possible thereafter.

There will be a limited number of PhD stipends not attached to specific projects..

In addition, there will be PhD stipends available in:

  • Actuarial science within the project frame InterAct (contact Mogens Steffensen,
  • Causality in event driven dynamical systems (contact Niels Richard Hansen,
  • Statistical and causal methodology for complex data structures with applications to microbiome data analysis (contact Niklas Pfister,
  • Topics on interpretable machine learning (contact Munir Hiabu,

You are welcome to contact any member of the department to discuss research projects.

Deadline : 01 April 2023

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(11) PhD fellowships in Mathematics

There will be PhD stipends available in:

  • Algebraic and Arithmetic Geometry, Applied algebra, Combinatorics, Geometric group theory, Geometry and Geometric Analysis, Number Theory, Representation Theory, and Topology as covered by the section Algebra & Geometryand  Copenhagen Center for Geometry & Topology (GeoTop).
  • Classical Analysis, Descriptive Set Theory, Functional Analysis, Geometric group theory, Harmonic Analysis, History of Mathematics, Mathematical Physics, Operator Algebras, Partial Differential Equations, Quantum Information, Quantum Computation and Quantum Simulation as covered by the section section Analysis and Quantum, the Centre for the Mathematics of Quantum Theoryfunded by the Villum Foundation and the ‘Quantum for Life‘ Center funded by the Novo Nordisk Foundation.

Deadline: 1  April 2023

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(12) PhD fellowship in Non-equilibrium quantum physics

The project aims to study emergence of complex dynamics in quantum many-body systems from the microscopic laws of quantum physics and will directly tie into various ongoing research directions in non-equilibrium quantum physics including dissipative and light-induced many-body engineering, quantum many-body non-ergodic dynamics (e.g. quantum many-body scars) and quantum information processing. Connection with experiments will be sought when possible. The project is based on unconventional algebraic principles discovered by the PI which are already finding broad applications ranging from operator algebras, through novel passive quantum error correcting algorithms in realistic quantum computing platforms to computing complex dynamics in driven quantum matter experiments. As a part of the PhD the student will have the opportunity to learn and contribute to developing new theoretical approaches to solving wide ranges of quantum many-body physics relevant to condensed matter physics, quantum computing and quantum science and technology, more broadly. 

Deadline : 31 March 2023

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(13) PhD fellowship in Quantum Photonics

The goal of the project is to develop a platform for photonic quantum information processing based on nano-opto-electro-mechanical systems (NOEMS) [1]. NBI has developed state-of-the-art deterministic single-photon sources [2] using semiconductor quantum dots integrated in photonic nanostructures. To fully harness the potential of integrated photonic quantum technologies, it is necessary to scale it up, i.e. to perform operations on many photonic quantum bits (or ‘qubits’) simultaneously to generate and manipulate entangled photons. The overarching goal is to demonstrate the seamless integration of single-photon sources with the quantum gates required for building truly scalable quantum photonic micro-processors. In this project we will pursue a novel approach to control light-matter interaction and quantum gates based exclusively on electro- and opto-mechanical interactions. The emphasis of this project will be in the design and fabrication of nanophotonic devices for scaling single-photon sources [3], with a focus on applications within quantum communication [4].

The PhD student’s tasks will include:

  • Learn and understand solid-state quantum emitters and the fundamentals of quantum optics (MSc and PhD level courses are offered at NBI).
  • Acquire the fabrication skills required to build nano-mechanical devices and single-photon sources in gallium arsenide (GaAs).
  • Perform optical characterization of nano-mechanical devices and photonic integrated circuits at cryogenic temperature.
  • Characterize the single-photon sources in our optical laboratories.
  • Perform numerical simulations (finite difference time domain or finite elements) of the devices.

Deadline : 26 March 2023

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(14) PhD scholarship in Fungal Evolution and Genomics

The Integrative Evolutionary Biology (IEB) group in the Department of Biology at the University of Copenhagen in Denmark is offering a PhD scholarship in fungal evolution and genomics starting 1 September 2023 or as soon as possible thereafter. The University of Copenhagen provides a top-notch research environment and the city of Copenhagen provides a vibrant cultural scene.

Project Description 

The PhD scholarship will be part of a newly established research project “Missing-link cultivars: using fungus-farming ants to solve mysteries of crop domestication” funded by the Carlsberg Foundation. More information about the project can be found here:

 The project will explore genomic signatures of crop domestication, focusing on a lineage of fungi that has been cultivated for food by attine ants for millions of years. These fungi have traits that reinforce their value as co-evolved food crops, and they depend on their ant farmers to provide them with nutritional substrates (e.g. fresh vegetation) foraged from the environment. The PhD candidate is expected to participate in the design of experiments and analyses to explore the cellular processes and metabolic pathways governing fungus crop performance. Important research approaches will include genomic, comparative genomic, bioinformatic, and experimental mycological techniques.

 Daily work will be based in the thriving research environment of the Section of Ecology and Evolution ( The project will involve a longer research stay abroad at a host institution to be determined.

Deadline :  20 March 2023

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(15) PhD fellowship in Sleep Dynamics at Center for Translational Neuromedicine

Sleep is essential for cognitive performance, and it is well established that impaired sleep reduces cognition. Sleep is not a homogeneous brain state but composed of several micro-structures regulated by many regions within the brain. In recent years, short arousals have been recognized as an integral part of normal sleep adding to the complexity of sleep. We are interested in determining the impact of these frequent sleep-arousal transitions on shaping restorative sleep processes related to both memory consolidation and waste clearance. Aging is characterized by sleep disturbances, which could contribute to age-related cognitive decline. In this project, the PhD student will investigate how alterations in sleep microarchitecture during aging affects fluid flow in the brain and explore novel therapeutic strategies related to regulation of sleep-arousal transitions.

Deadline : 20 March 2023

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(16) PhD position: ERC-Funded PhD Project in Mechanobiology and Active Matter

Active matter comprises a wide range of living systems including cells, bacteria, and subcellular filaments, as well as artificial and synthetic materials that are continuously driven far from equilibrium by their constituent elements. Fundamental biological processes such as cell division, cell death, and the emergence of coordinated motion of cells are governed by the activation of biochemical signals by mechanical stresses and by the physics of active matter. This research program will use theoretical and computational modeling with potential applications in the life sciences.

Who are we looking for?

We are looking for candidates within the field(s) of soft matter physics. Applicants can have a background from theoretical and computational modeling of soft materials.

Our group and research- and what do we offer?

At the Active Intelligent Matter Group we combine multiscale modeling – discrete and continuum simulations – with in-house experiments to address various aspects of complexities in active materials. This spans a wide range of problems from revealing the impact of mechanical forces on biochemical signalling in cells, investigating the dynamics of bacterial competition, to design and control of bio-inspired materials that are capable of self-organization, self-healing, and self-pumping.

We offer a highly creative and collaborative research atmosphere at the interface between physics, biology and fluid mechanics. The PhD fellowship is given in accordance with Danish legislation and will typically run for 3 years. The salary is set accordingly to Danish Union Contracts which are quite competitive on an international scale.

The group is a part of Niels Bohr Institute, Faculty of SCIENCE, University of Copenhagen. We are located in Copenhagen.

We offer creative and stimulating working conditions in dynamic and international research environment. Our research facilities include modern laboratories and high performance computing facilities.

Deadline: 20 March 2023

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